A coherent cosmic background of axion-like particles (ALPs) coupled to photons can produce a small periodic differential phase or polarization modulation for photons traversing separate arms in gravitational wave interferometers, peaked at a frequency associated with the particle mass, and suppressed whenever the dark-matter coherence length exceeds the arm length~. For the LIGO audio frequency band the sensitivity to cosmic ALPs is below current bounds. For LISA, however, the natural mass range –~eV – corresponding to a sideband frequency in LISA’s science band of ~mHz–~Hz – may be observable. A 1-year shot-noise-limited search projects a sensitivity ~GeV across most of the band, reaching ~GeV near ~Hz, which is – below the CAST helioscope bound. An RF heterodyne photodetection upgrade – to either detector – might extend the search sensitivity to ~GeV at ~eV for LIGO and ~GeV at ~eV for LISA. Dark-matter substructure can affect the signal in a several interesting ways.

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