Let be the moduli space of hyperbolic surfaces of genus endowed with the Weil-Petersson metric. In this paper, we show that for any , as , for a generic surface in , the error term in the Prime Geodesic Theorem is bounded from above by , up to a uniform constant multiplication. The expected value of the error term in the Prime Geodesic Theorem over is also studied. As an application, we show that as , on a generic hyperbolic surface in most closed geodesics of length significantly less than are simple and non-separating, and most closed geodesics of length significantly greater than are not simple, which confirms a conjecture of Lipnowski-Wright. A novel effective upper bound for intersection numbers on is also established, when certain indices are large compared to .

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