Capture-recapture experiments are widely used to estimate the abundance of a finite population. Based on capture-recapture data, the empirical likelihood (EL) method has been shown to outperform the conventional conditional likelihood (CL) method. However, the current literature on EL abundance estimation ignores behavioral effects, and the EL estimates may not be stable, especially when the capture probability is low. We make three contributions in this paper. First, we extend the EL method to capture-recapture models that account for behavioral effects. Second, to overcome the instability of the EL method, we propose a penalized EL (PEL) estimation method that penalizes large abundance values. We then investigate the asymptotics of the maximum PEL estimator and the PEL ratio statistic. Third, we develop standard expectation-maximization (EM) algorithms for PEL to improve its practical performance. The EM algorithm is also applicable to EL and CL with slight modifications. Our simulation and a real-world data analysis demonstrate that the PEL method successfully overcomes the instability of the EL method and the proposed EM algorithm produces more reliable results than existing optimization algorithms.

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