We describe the Galois action on the middle -adic cohomology of smooth, projective fourfolds that occur as a fiber of the Albanese morphism on moduli spaces of sheaves on an abelian surface with Mukai vector . We show this action is determined by the action on and on a subgroup , which depends on . This generalizes the analysis carried out by Hassett and Tschinkel [HT13] over . As a consequence, over number fields, we give a condition under which and are not derived equivalent. The points of correspond to involutions of . Over , they are known to be symplectic and contained in the kernel of the map . We describe this kernel for all varieties of dimension at least . When is a fourfold over a field of characteristic 0, the fixed-point loci of the involutions contain K3 surfaces whose cycle classes span a large portion of the middle cohomology. We examine the fixed loci in fourfolds over where is a -polarization, finding the K3 surface to be elliptically fibered under a Lagrangian fibration of .

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