For a connected reductive group and an affine smooth -variety over the complex numbers, the localization functor takes -modules to -modules. We extend this construction to an equivariant and derived setting using the formalism of h-complexes due to Beilinson-Ginzburg, and show that the localizations of Harish-Chandra -modules onto have regular holonomic cohomologies when are both spherical reductive subgroups. The relative Lie algebra homologies and -branching spaces for -modules are interpreted geometrically in terms of equivariant derived localizations. As direct consequences, we show that they are finite-dimensional under the same assumptions, and relate Euler-Poincaré characteristics to local index theorem; this recovers parts of the recent results of M. Kitagawa. Examples and discussions on the relation to Schwartz homologies are also included.

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