Let or and regarded as a subgroup of . Here, is the quaternion division algebra over . For a character on , we say that an irreducible smooth admissible moderate growth representation of is -distinguished if . We compute the root number of a -distinguished representation twisted by the representation induced from . This proves an Archimedean analogue of the conjecture by Prasad and Takloo-Bighash (J. Reine Angew. Math., 2011). The proof is based on the analysis of the contribution of -orbits in a flag manifold of to the Schwartz homology of principal series representations. A large part of the argument is developed for general real reductive groups of inner type. In particular, we prove that the Schwartz homology is finite-dimensional and hence it is Hausdorff for a reductive symmetric pair and a finite-dimensional representation of .

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