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In this paper, we introduce tree varieties as a natural generalization of products of partial flag varieties. We study orbits of the PGL action on tree varieties. We characterize tree varieties with finitely many PGL orbits, generalizing a celebrated theorem of Magyar, Weyman and Zelevinsky. We give criteria that guarantee that a tree variety has a dense PGL orbit and provide many examples of tree varieties that do not have dense PGL orbits. We show that a triple of two-step flag varieties has a dense PGL orbit if and only if .

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