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Let S be a K3 surface and Aut D(S) the group of auto-equivalences of the derived category of S. We construct a natural representation of Aut D(S) on the cohomology of all moduli spaces of stable sheaves (with primitive Mukai vectors) on S. The main result of this paper is the precise relation of this action with the monodromy of the Hilbert schemes S^[n] of points on the surface. A formula is provided for the monodromy representation, in terms of the Chern character of the universal sheaf. Isometries of the second cohomology of S^[n] are lifted, via this formula, to monodromy operators of the whole cohomology ring of S^[n].

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