The production of gravitational vacuum defects and their contribution in energy density of the Universe are discussed. These topological microstructures could be produced as the result of defect creation of the Universe from “nothing” as well as the result of the first relativistic phase transition. They must be isotropically distributed on background of the expanding Universe. After Universe inflation these microdefects smoothed, stretched and broke up. Parts of them have survived and now they are perceived as the structures of Lambda-term (quintessence) and unclustered dark matter. It is shown that for phenomenological description of vacuum topological defects of different dimensions (worm-holes, micromembranes, microstrings and monopoles) the parametrizational noninvariant members of Wheeler -DeWitt equation can be used. The mathematical illustration of these processes may be the spontaneous breaking of local Lorentz-invariance of quasi-classical equations of gravity. In addition, 3-dimensional topological defects revalues Lambda-term.

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