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The ratio of the muon flux density to charged particle flux density at distances of 300 and 600 m from the shower axis ( and ) is measured. In addition, the energy dependence of is analysed for showers with energies above eV. A comparison between the experimental data and calculations performed with the QGSJET model is given for the cases of primary proton, iron nucleus and gamma- ray. We conclude that the showers with eV can be formed by light nuclei with a pronounced fraction of protons and helium nuclei. It is not excluded however that a small part of showers with energies above eV could be initiated by primary gamma-rays.

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