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In this paper, we conduct the first systematic investigation of twisted cubics on Gushel-Mukai (GM) fourfolds. We then study the double EPW cube, a 6-dimensional hyperkähler manifold associated with a general GM fourfold , through the Bridgeland moduli space, and show that it is the maximal rationally connected (MRC) quotient of the Hilbert scheme of twisted cubics on . We also prove that a general double EPW cube admits a covering by Lagrangian subvarieties constructed from the Hilbert schemes of twisted cubics on GM threefolds, which provides a new example for a conjecture of O’Grady.

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