We establish metric graph counterparts of Pleijel’s theorem on the asymptotics of the number of nodal domains of the -th eigenfunction(s) of a broad class of operators on compact metric graphs, including Schr"odinger operators with -potentials and a variety of vertex conditions as well as the -Laplacian with natural vertex conditions, and without any assumptions on the lengths of the edges, the topology of the graph, or the behaviour of the eigenfunctions at the vertices. {Among other things, these results characterise the accumulation points of the sequence , which are shown always to form a finite subset of . This} extends the previously known result that , for certain realisations of the Laplacian, in several directions. In particular, in the special cases of the Laplacian with natural conditions, we show that for graphs with rationally dependent edge lengths, one can find eigenfunctions thereon for which ; but in this case even the set of points of accumulation may depend on the choice of eigenbasis.

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