Zeros of two-dimensional sums of the Epstein zeta type over rectangular lattices of the type investigated by Hejhal and Bombieri in 1987 are considered, and in particular a sum first studied by Potter and Titchmarsh in 1935. These latter proved several properties of the zeros of sums over the rectangular lattice, and commented on the fact that a particular sum had zeros off the critical line. The behaviour of one such zero is investigated as a function of the ratio of the periods of the rectangular lattice, and it is shown that it evolves continuously along a trajectory which approaches the critical line, reaching it at a point which is a second-order zero of the rectangular lattice sum. It is further shown that ranges of the period ratio can be so identified for which zeros of the rectangular lattice sum lie off the critical line.

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