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Period domains, the classifying spaces for (pure, polarized) Hodge structures, and more generally Mumford-Tate domains, arise as open –orbits in flag varieties . We investigate Hodge–theoretic aspects of the geometry and representation theory associated with these flag varieties. In particular, we relate the Griffiths–Yukawa coupling to the variety of lines on (under a minimal homogeneous embedding), construct a large class of polarized –orbits in , and compute the associated Hodge–theoretic boundary components. An emphasis is placed throughout on adjoint flag varieties and the corresponding families of Hodge structures of levels two and four.

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