We establish the dynamical attractor behavior in scalar-tensor theories of dark energy, providing a powerful framework to analyze classes of theories, predicting common evolutionary characteristics that can be compared against cosmological constraints. In the Jordan frame the theories are viewed as a coupling between a scalar field, \Phi, and the Ricci scalar, R, F(\Phi)R. The Jordan frame evolution is described in terms of dynamical variables m \equiv d\ln F/d\ln \Phi and r \equiv -\Phi F/f, where F(\Phi) = d f(\Phi)/d\Phi. The evolution can be alternatively viewed in the Einstein frame as a general coupling between scalar dark energy and matter, \beta. We present a complete, consistent picture of evolution in the Einstein and Jordan frames and consider the conditions on the form of the coupling F and \beta required to give the observed cold dark matter (CDM) dominated era that transitions into a late time accelerative phase, including transitory accelerative eras that have not previously been investigated. We find five classes of evolutionary behavior of which four are qualitatively similar to those for f® theories (which have \beta=1/2). The fifth class exists only for |\beta| < \sqrt{3}/4, i.e. not for f® theories. In models giving transitory late time acceleration, we find a viable accelerative region of the (r,m) plane accessible to scalar-tensor theories with any coupling, \beta (at least in the range |\beta| \leq 1/2, which we study in detail), and an additional region open only to theories with |\beta| < \sqrt{3}/4.

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