Higher-Order Competition as a Minimal Mechanism for Spatial Pattern Diversity in Population Dynamics

We show that negative feedback alone generates the diversity of self-organized shapes usually attributed to scale-dependent activation-inhibition. For a broad class of kernels, pairwise competition models only generate hexagonal spot arrays. Higher-order terms eliminate this restriction and promote stripes and gaps. Combining individual-based simulations and nonlinear analysis, we derive the pattern-selection thresholds and construct the full state diagram, including an unusual spots-stripes-spots sequence. Spot patterns are thus not a reliable indicator of proximity to a tipping point.