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We study the structure of cyclic Riemannian nilmanifolds, that is, connected nilpotent Lie groups endowed with a left-invariant cyclic metric . We prove that a Riemannian nilmanifold is cyclic if and only if its Lie algebra is at most two-step nilpotent and the associated family of skew-symmetric endomorphisms is Abelian. As a direct consequence, every cyclic Riemannian nilmanifold is naturally reductive. We also determine the full isometry group of a connected and simply connected cyclic Riemannian nilmanifold without Euclidean factor.

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